Tuesday, 10 January 2017

Jean Baptiste Pierre Antoine de Monet, Chevalier de Lamarck

السلام عليكم و رحمة الله

This name (in the title) sounds like some commander or soldier from the past and you're right. Jean Baptiste Lamarck is a truly forgotten man of our time, in fact he's only known by the "long giraffe neck eating leaves from trees" man. Even Subboor, in his recent debate, mentioned that. But far from that, Lamarck was actually quite different from who we think he is or what our textbooks say about him.

Jean Baptiste Lamarck
Let me be frank, I can not speak French but I have read a bit of his translated work about his hypothesis and I noticed the obvious connection between Lamarck and Charles Darwin.

Lamarck was born on 1st August 1744, long before Darwin. Later in his life, Lamarck devised his hypothesis of how evolution occurred. His hypothesis was rejected despite his fame on the work on invertebrates (he coined that word -  so blame him if you hated that word in your high school). It was rejected so much that he was rejected by people and he died blind with his 2 daughters serving him in his old age. He published his hypothesis in the month of "flowering"[1] but I guess he didn't get a flowery end.

Lamarck was born in a family that was devoted to military but Lamarck was sent, by his father, to Jesuit College for ecclesiastical (relating to church) career. He never liked it. After his father died, Jean left the Jesuit College and became a soldier to fight in the battle of Fissingshausen - but lost. His fellow officers were all killed. Later on he stopped engaging in war and went to Monaco (very little tiny place next to France where rich people live and pay no tax - so called "equality") and in Monaco, his comrade physically lifted Lamarck from his head! What a fool. His lymphatic glands (in the neck) caught inflammation and thus he was forced to leave his position as a lieutenant (which he had been promoted to).  After this, due to his health not being good, he worked in a bank. He then took a course in medicine and met Jean-Jacques Rousseau. Lamarck preferred, in fact loved, botany than medicine and so he left medicine for it. Anyway, in 1781, Lamarck received a commission to visit botanical gardens and institutions in Germany, Hungary and Holland. He collected rare plants and made notes for Jardin du Roi botanical garden in Paris. Then, Museum d'Histoire Naturelle was founded. Lamarck was the chairman of zoology in Naturelle. As a chairman, he threw botany to one side and studied invertebrates and published 7 volumes on them in his Histoire Naturelle des Animaux sans vertebres (1815-1822). He also published a book on meteorology but nothing important was in it and he published Philosophie Zoologique. With his meteorology book, which, according to the people at the time contained ridiculous speculation and so he was ridiculed at the time. But with his Zoologique book which is based on natural history, he went to the emperor Napoleon and handed him his book (perhaps Lamarck's motive in giving Napoleon his book was to get his fame back since he was ridiculed?). However Napoleon said:

"What is this? said the Emperor. "Is it your absurd Meteorologie with which you are disgracing your old age? Write on natural history, and I will receive your works with pleasure. This volume I only accept out of consideration for your grey hair. Here"[2] (Napoleon stated as he handed the book to someone else, perhaps his servant).

The book was about natural history but Lamarck didn't say that, he just cried.

Today Lamarck's hypothesis, is more or less forgotten, but it was much more than just inherited acquired characteristic, which simply means, whatever a person acquires during his life time, like, bigger muscles, this acquired characteristic is inherited to his descendants; thus the muscles of the children would improve. Or the famous giraffe neck. Giraffes stretch their necks, this characteristics is inherited to other giraffes and eventually over long period of time, giraffes have long necks.His hypothesis had two main principles: (1) inherent progress and (2) adaptation in environment. That is for life after it appearance. So where did life itself come from? Lamarck believed that it naturally appeared from molecules,

"Life and organisation are products of nature, and at the same time results of the powers conferred upon nature by the Supreme Author of all things and of the laws by which she herself is constituted: this can no longer be called in question. Life and organisation are thus purely natural phenomena, and their destruction in any individual is also a natural phenomenon, necessarily following from the first"[3]

Inherent progress


Ornithorhyncus (or Platypus).
An animal with "duck mouth"(a bird
characteristic) with a body of a mammal
but also lays eggs whilst mammals,
like humans, don't lay eggs.
Perfect creation, a wonder to ponder over.
Lamarck believed that not only life is spontaneous (life from non-life) but after life's appearance, it naturally evolves, hence, inherent progress. Thus, because life is constantly evolving from non molecules, it means that there is no such thing as extinction since life according to Lamarck is like a ladder and there is always a creature evolving to the new stage. So imagine a ladder with 10 steps, and life steps on the first step and then another life appears spontaneously and also steps on the first step of the ladder but by the time the second life hits the first step, the first life is already on the second step of the ladder and so on. Thus each step is occupied and therefore no such thing as extinction occurs. So why do we have a clear cut distinction between, say mammals and birds? Such a big gap must not exist. Lamarck tried to solve this by pointing to the fact that we still haven't discovered or identified all creatures on Earth. But when we find them, then we can fill the gaps. But Lamarck didn't just blame the lack of discovery but he postulated an example which "proves" his hypothesis. He used the Ornithorhyncus (Platypus) as proof that here is a mammal with a characteristic of a bird. Lamarck called them (with Echidna) "Animals intermediate between birds and mammals"[4]
Echidna.
Also a mammal but lays eggs.

Adaptation in environment

Lamarck believed that animals can also change due to the environment they live in. The reason why a giraffe has a long neck he argued was due to the fact that trees are a little long, and so the giraffes must stretch their necks in order to reach the trees (couldn't the giraffe just eat from the ground?) and therefore this stretch is gradually increasing the length of the neck by the mechanics that he postulated and called inherent acquired characteristics. Due to the fact that the environment in which the animal resides in - the adaptation changes slowly too, it logically means that animals too change slowly,

"Not only is this the greatest marvel that the power of nature has attained, but it is besides a proof of the lapse of a considerable time; since nature has done nothing but by slow degrees"[5]

Depending where you live, that would, in a Lamarckian worldview, determine whether you will use certain body parts or not. This is where his "law of use and disuse" comes in. So if a fish lived in a cave which is dark, the fish won't use it's eyes, thus based on the law of use and disuse, the eyes will eventually disappear in the future generations since it's not using the eye. Which is why Lamarck said that environment (and therefore the mode of life) controls the body.[6] Lamarck used the eyes of the mole as an example. He said "Yet the mole, whose habits require a very small use of sight, has only minute and hardly visible eyes, because it uses that organ so little"[7]

Even though this is a very summarized explanation of Lamarck's hypothesis, you can, nevertheless see gradualism, adaptation and evolution in it, which is from where Charles Darwin got some of his ideas from. Of course Darwin got his ideas from others too but Lamarck's works no doubt influenced Darwin, especially when Darwin read Lamarck's work via Charles Lyell's critique of Lamarckism in Principles of Geology volume 2 and owning Philosophical Zoology.[8]

By comparing Lamarck's evolutionary ideas and Darwin's evolutionary ideas we see very clear cut similarities and the differences between them are fairly small. Both accepted gradualism (geology and biology), evolution, environmental adaptation and the law of use & disuse. The difference is that according to Lamarck, environmental adaptation is a force that is second to the inherent progress whilst Darwin postulated adaptation as the force itself that made all the diversity. Lamarck accepted inherent progress whilst Darwin accepted adaptation as the factor that causes evolution, so Darwin rejected inherent progress.

Summary

There is no doubt that Lamarck's ideas are very similar to Darwin's and therefore I conclude that Lamarck was one of the factors that influenced Darwin. Of course Robert Chambers & Erasmus Darwin and other evolutionists influenced Darwin as well but it seems clear that Lamarck influenced him a great deal.

Note: Yes Lamarck did use the giraffe as an example, that was not made up. Lamarck said,

"It is interesting to observe the result of habit in the peculiar shape and size of the giraffe (Camelo-pardalis): this animal, the largest of the mammals, is known to live in the interior of Africa in places where the soil is nearly always arid and barren, so that it is obliged to browse on the leaves of the trees and to make constant effort to reach them. From this habit long maintained in all its race, it has resulted that the animal's fore-legs have become longer than its hind legs, and that its neck lengthened to such a degree that the giraffe, without standing up on its hind legs, attains a height of six meters (nearly 20 feet)"[9]

That's the only time he spoke about giraffes. He spoke about adaptation much more, in fact a whole chapter is dedicated to it called "Of the influence of the environment on the activities and habits of animals",  but giraffes are the only thing we know of his hypothesis. I guess Lamarck answered my question "couldn't the giraffe just eat from the ground?" when he said "it is obliged to browse on the leaves of the trees and to make constant effort to reach them" but what about the animals that don't have long necks? They still coexist with giraffes...

The cause of his blindness (for 10 years until he died) was suggested to be due to his hard work involving looking through the microscope too much - a legend that deserves respect.

و الله اعلم


References

[1] Stephen Jay Gould, 2002, The Structure of Evolutionary Theory. Belknap Harvard, page 170 and Stephen Jay Gould, 2001, The Lying Stones of Marrakech. Vintage, page 115.

[2] Translation by Hugh Elliot of Lamarck's Philosophie Zoologique. Hugh Elliot, 1963, Zoological Philosophy: An exposition with regard to the natural history of animals, Hafner Publishing Company, page xxi.

[3] Ibid, page 236.

[4] Ibid, page 166.

[5] Ibid, page 50.

[6] Ibid, page 106.

[7] Ibid, page 116.

[8] Stephen Jay Gould, 2002, The Structure of Evolutionary Theory. Belknap Harvard, page 195

[9] Hugh Elliot, 1963, Zoological Philosophy: An exposition with regard to the natural history of animals, Hafner Publishing Company, page 122

Friday, 6 January 2017

Stephen Jay Gould

السلام عليكم و رحمة الله

Stephen Jay Gould (1941-2002) is, without a doubt, a forgotten legend among the evolutionists of today because of their preference to gradualism. He is remembered just for his punctuated equilibrium hypothesis which was designed to challenged two main aspects of the conventional wisdom of the time, gradualism (on macro-evolutionary level) and individual selection[1].

But his general work ranging from classical literature to modern literature (that would be 1970-2000 in his time) is forgotten and I decided to write this article to remember some, in fact very little (when you look at the huge number and size of his books) of his work and it's importance to the overall landscape of evolutionary debate.

Stephen Jay Gould in his office

One of my favorite things of Gould is his Cordelia dilemma[2]. He named it 'Cordelia' to remember the terrible fate of King Lear who had 3 daughters. Lear wanted to leave the biggest share of his kingdom, after his death, to the daughter that professed the most love to him. So two of the daughters, Goneril & Regan, excessively lied about their love to him and Cordelia simply remained silent, or hardly said a thing because she believed that the love in her heart was more "ponderous" (i.e. large) than the tongue. Lear got angry and disowned her which affected him and lead to his madness, blindness and death.[3] But Gould used her as an example because nature might 'say' something whilst we don't recognize it - in fact we don't want to recognize it because of the a priori belief in gradualism. What Gould meant was that the conventional Neo-Darwinian belief in gradualism was not supported by the fossil record, rather because Charles Darwin (1809-1882) wanted extreme slowness and this was simply adopted by the predecessors evolutionist after Darwin.

When Darwin published his Origin of Species (1859) he made it clear that evolution works in a very gradual and slow manner, depending on the rate of birth but nevertheless slow & gradual. For example Darwin said,

"Although each formation may mark a very long lapse of years, each perhaps is short compared with the period requisite to change one species into another"[4]

"Nature acts uniformly and slowly during vast periods of time..."[5]

And since the formulation of Darwinism (1859-1890's), which advocated gradualism, Neo Darwinism (1890's - 1970's) came out with the same gradualistic belief. This idea of gradualism advocated the view that evolution is very slow, bit by bit and therefore we should look at the fossil record and see no same creature in two completely different ages. So let's say in zone A we see a fish, we expect it to be gone by the time it reaches zone B (assume that the time between zone A and B are very long, say a million years) but if we see a fish in zone Z then clearly that fish survived and did not change whatsoever from zone A all the way to zone Z (that would be 25 million years of no change). So if for such a long time, we do not see gradual change but rather stability then you cannot mix the two ideas together. So how did the Neo Darwinist reconcile such contradictory results between the theoretical, a priori, assumption and stability? Gould shares 2 main important points:

1. It wasn't shared in the technical literature.

2. If it was known, it would've been seen as "negative" result.

It wasn't shared in the technical literature.

Gould was aghast by the fact that not many biologists know the fact that stasis ruled the paleontological world. He said,

"The common knowledge[6] of a profession often goes unrecorded in technical literature for two reasons: one need not preach commonplaces to the initiated; and one should not attempt to inform uninitiated in publications they do not read. The longterm stasis, following a geologically abrupt origin of most fossil morphospecies, has always been recognized by professional paleontologists..."[7]

So the reasons why it was not shared are because; you do not have to repeat the same fact known by many already and no need to teach the laymen, like us, since we do not read such publication. Such knowledge which surely does question the Neo darwinian mechanism was held back for the sake of a theoretical belief.

If it was known, it would've been seen as "negative" result.


Brachiopod fossil. 
My personal collection. 
From Carboniferous rocks. 
(300+ million years old)
It's still alive today. 
Gould shares a personal story of his personal Ph.D adviser John Imbrie. Imbrie was a paleoclimatologist who accepted gradualism because he simply was taught it. But Imbrie went out and studied 30 different species of brachiopods from the Devonian era (in Michigan Basin) and realized that all but one have remained stable for all those millions upon millions of years, that other one was ambiguous. But when he shared his results, it wasn't published as a...

"...triumphant paper documenting the important phenomenon of stasis. Instead, he just become disappointed at such "negative" results after so much effort. He buried his data in a technical taxonomic monograph that no working biologist would ever encounter... and eventually left the profession for something more "productive""[8]

Such problems have no doubt happened again and again and the reason why stasis was seen as "negative" result was because stasis was not expected. Gould, on the other hand, does not see the stasis as a negative evidence but rather as positive evidence that the fossil record is signalling (obviously). The reason why stasis was seen as negative was also because all evidence, no matter which one, is always interpreted through a theory. Evidence doesn't speak. You interpret the evidence. That's another reason why stasis was ignored even though known by many paleontologists.
Bivalves. 
Personal collection. 
Carboniferous rocks.
Alive today.

That is Cordelia's dilemma. Stasis is data![9]

Another thing I like about Gould is his Petrus Camper story[10].

Petrus Camper (1722-1789) was a Dutch physician who was intrigued by what 'methods' the Greeks used to make sculptures of their gods for the face. He claimed to have discovered it and published an image in his thesis (see images below). Camper never intended racism but you can easily see how racism is extrapolated from his work.










In the Academy of Drawing in 1770 the director distorted Camper's view and told the people that these evidences were evidence for the Great Chain of Being[11] (an evolutionary idea which promoted inherent progress evolution and that the higher you are on the scale, the better evolved you are, the lower you are on the scale, the less evolved you are). What is strange is that Petrus was a monogenist (believed in a single origin for mankind) yet his work was used by polygenist! It's amazing how ideas get circulated around to racist people like Jules J. Virey, Samuel T. Soemmering and no doubt to the hands of the Nazis & to many eugenicist (inspired by Social Darwinism which was inspired by Charles Darwin from both his Origin of Species and Descent of Man).

Scala Natura (The scale of nature)
Beware of distorted evidence.

But as Gould ends his essay on Petrus, he said,

"He became the semiofficial grandpappy of the quantitative approach to scientific racism, yet his own concept of human variability precluded judgements about innate worth a priori... He became a villain of science when he tried to establish criteria for art. Camper got a bad posthumous shake on earth..."[12]

What amazes me is that Gould actually knew this stuff. Seldom do you find an evolutionist who is open to such things (at least on a public level) even though Gould was silent on the racism perpetuated by Charles Darwin.

The final thing I like about Gould is his timeless critique of Neo Darwinism.

Despite the strong wall built by Neo Darwinist since 1959, Gould simply did not buy their theory. Gould's legacy, perhaps the most important legacy is his critique of Neo Darwinism. You see, ideas are sometimes taught as fact, like Neo Darwinism, to kids. Due to such indoctrination, those kids, when they grow up, won't really know the difference between fact from fiction and simply accept that "smart" scientist with white coats and goggles know their stuff even though they themselves understand nothing. Anagenesis, cladogenesis, homology, convergent evolution... have you heard of them? No. Do you understand them? No. Actually you have and do but those words are there to fool you so you simply accept that "smart" guys know it all.

Every hypothesis has it's core and for Neo Darwinism there are 3 main cores: gradualism, individual selection and microevolution.

Gould (as I have shown in the first point) critiqued gradualism because the fossil record doesn't fit the hypothesis of Neo Darwinism and he critiqued microevolution to macroevolution. Microevolution is a flimsy word and generally it's a trap but there is a definite limit between micro and macroevolution. He didn't critique it and left it unhealed but rather he used other causes that he thought were important to cause macroevolution. These causes are known among many evolutionist today[13] The reason why Gould critiqued the microevolution to macroevolution claim was because if it's true then we expect gradualism than stasis and also we don't expect to see abrupt appearance in the fossil record. Abrupt appearance is the sudden appearance of a creature. The creature(s) just "pops" into existence without any prior ancestor. Due to this discontinuous pattern, micro to macro cannot possibly be the explanation and that is why Gould critiqued it. But apart from Gould, there are other evidences as well that perhaps would've impressed Gould (actually refuted evolution altogether). It turns out that one protein cannot evolve into another protein even if they are similar.[14] That means evolution couldn't have possibly occurred since we have many proteins and other cellular machines. Another point is that developmental gene regulatory networks cannot be mutated haphazardly either. If you introduce a change to it, it will cause death to the growing embryo. That is why Eric Davidson (who did the experiments on the developmental gene regulatory networks) said,

"Neo-Darwinian evolution is uniformitarian in that it assumes that all process works the same way, so that evolution of enzymes or flower colors can be used as current proxies for study of evolution of the body plan. It erroneously assumes that change in protein coding sequence is the basic cause of change in developmental program; and it erroneously assumes that evolutionary change in body plan morphology occurs by a continuous process. All of these assumptions are basically counterfactual. This cannot be surprising, since the neo-Darwinian synthesis from which these ideas stem was a pre-molecular biology concoction focused on population genetics and adaptation natural history, neither of which have any direct mechanistic import for the genomic regulatory systems that drive embryonic development of the body plan"[15]

Not only does the fossil record fail to show what Neo Darwinist need but recent evidence is building up to show that it can't happen which is why, in the recent Royal Society meeting held in London, did evolutionary scientists from across the world meet to discuss these problems facing the orthodox evolution taught in schools.

Anyway, Gould left a lot of knowledge but I have only shared a little.

By the way, do not think I believe in punctuated equilibrium that Gould proposed, I don't accept it. However if someone says what is right or interesting then it must be known.

و الله اعلم


Reference

1. Stephen J. Gould, 2002, The Structure of Evolutionary Theory. Belknap Harvard.

2. Ibid and Stephen J. Gould, 1995, Dinosaur in a Haystack. Harmony Books.

3. It is a Shakespeare play King Lear. I have not heard of it prior to Gould.

4. Darwin: The Origin of Species, Wordsworth Classics of World Literate, page 222.

5. Ibid, page 205

6. Common knowledge means knowledge known by most people. And the common knowledge in this case is stability, or stasis.

7. Stephen J. Gould, 2002, The Structure of Evolutionary Theory. Belknap Harvard, pages 749-750

8. Ibid, page 760.

9. Ibid, page 759.

10. Stephen Jay Gould, 1991, Bully for Brontosaurus. Hutchinson Radius.

11. Gustav Jahoda, 1999, Images of Savages: Ancient Roots of Modern Prejudice in Western Culture, Routledge, page 73.

12. Stephen Jay Gould, 1991, Bully for Brontosaurus. Hutchinson Radius, page 240.

13. Douglas H. Erwin, 2000, Macroevolution is more than repeated rounds of microevolution, Evolution & Development, 2:2, 78-84,

14. Ann Gauger and Douglas Axe. The Evolutionary Accessibility of New Enzymes Functions: A Case Study from the Biotin Pathway. Bio-Complexity.

15. Eric Davidson, 2011, Evolutionary Bioscience as regulatory systems biology. Developmental Biology, volume 357, issue 1, 1 September, pages 35-40.